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The International Conference on Life System Modeling and Simulation (LSMS) was formed to bring together international researchers and practitioners in the field of life system modeling and simulation as well as life system-inspired theory and methodology. The arrival of the 21st century has been marked by a resurgence of research interest both in arriving at a systems-level und- standing of biology and in applying such knowledge in complex real-world appli- tions. Consequently, computational methods and intelligence in systems, biology, as well as bio-inspired computational intelligence, have emerged as key drivers for new computational methods. For this reason papers dealing with theory, techniques and real-world applications relating to these two themes were especially solicited.